Are you talking about the studs that hold the wheels on (the ones with the "R" stamp in your pictures)? Or the ones holding the locking mechanism into the holes in the picture?
I've never removed the wheel studs, but I think they should just be pressed in.
On the four front hubs I've disassembled, the locking mechanism was held on by bolts, not studs. And I didn't have any problems removing them.
On the newer axle I took apart, though, there was a Spicer type front axle shaft, and it has a C-clip at the end. On that one, I had to disassemble the locking mechanism itself to be able to access and remove the clip. But once I did that, everything came apart easily.
-Paul